Where did Herbert Boyer complete his graduate work?

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Herbert Boyer was born in 1936 and brought up in a corner of western Pennsylvania where railroads and mines were the destiny of most young men . He completed graduate work at the University of Pittsburgh, in 1963 , followed by three years of post-graduate studies at Yale. In 1966, Boyer took over assistant professorship at the University of California at San Francisco. By 1969, he performed studies on a couple of restriction enzymes of the E. coli bacterium with especially useful properties. Boyer observed that the